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The website was founded by Justin Sevakis in July 1998.
In May 2000, CEO Christopher Macdonald joined the website editorial staff, replacing editor-in-chief Isaac Alexander.
In July 2002, Anime News Network launched its Encyclopedia, a collaborative database of anime and manga titles also including information about the staff, cast, and companies involved in the production or localization of those titles.
On September 7, 2004, the Sci Fi Channel online newsletter Sci Fi Weekly named the site the Web Site of the Week.
In January 2007, ANN launched a separate version for Australian audiences.
On July 4, 2008, ANN launched its video platform with a library of anime trailers as well as its own news show ANNtv.
On August 7, 2017, a hacker took control of Anime News Network's domain (animenewsnetwork.com), and compromised some of the site's Twitter accounts, including the personal accounts of ANN's CEO Christopher Macdonald and Executive Editor Zac Bertschy.
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