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In bacterial pili, an ATPase hexamer rotates at the base of the pilus, which is responsible for pili extension and retraction (Chang et al., 2016; McCallum, Tammam, Khan, Burrows, & Howell, 2017). This rotational property appears to have been adapted in archaella to be used for swimming. It is interesting that the Archaea swimming mode resembles the bacterial flagella swimming mode (Kinosita, Uchida, Nakane, & Nishizaka, 2016), because it likely evolved from pili that display a different mode of motility.
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